Dr Henry Ruhl

Associate Head, Ocean Biogeochemistry and Ecosystems
Ocean Biogeochemistry and Ecosystems
+44 (0)23 8059 6365

I specialize in biological carbon cycle processes and ecology with a focus on the deep sea. The importance of describing such natural phenomena is growing as energy, fisheries, and mining activities continue to develop in deeper waters. Understanding contemporary processes will be crucial in evaluating any anthropogenic impacts in the deep-ocean. I am currently the PI for the NERC Project: Autonomous Ecological Surveying of the Abyss. In addition to deep-sea ecological research I’m helping to coordinate ocean observatory efforts related to EMSO, FixO3, DELOS and other efforts. These programs work not only through government funded research, but also by collaborating with industry. Such objective collaboration is already leading to outcomes not otherwise possible through academic or industrial research alone.



Durden, J.M..; Ruhl, H.A.; Pebody, C.; Blackbird, S.J.; van Oevelen, D.. 2017 Differences in the carbon flows in the benthic food webs of abyssal hill and plain habitats. Limnology and Oceanography. 10.1002/lno.10532 (In Press)
Yool, Andrew; Martin, Adrian P.; Anderson, Thomas R.; Bett, Brian J.; Jones, Daniel O.B.; Ruhl, Henry A.. 2017 Big in the benthos: future change of seafloor community biomass in a global, body size-resolved model. Global Change Biology. 10.1111/gcb.13680 (In Press)
Main, C.E.; Yool, A.; Holliday, N.P.; Popova, E.E.; Jones, D.O.B.; Ruhl, H.A.. 2017 Simulating pathways of subsurface oil in the Faroe–Shetland Channel using an ocean general circulation model. Marine Pollution Bulletin, 114 (1). 315-326.10.1016/j.marpolbul.2016.09.041
Milligan, R.J.; Morris, K.J.; Bett, B.J.; Durden, J.M.; Jones, D.O.B.; Robert, K.; Ruhl, H.A.; Bailey, D.M.. 2016 High resolution study of the spatial distributions of abyssal fishes by autonomous underwater vehicle. Scientific Reports, 6. 26095.10.1038/srep26095
Durden, Jennifer M.; Bett, Brian J.; Schoening, Timm; Morris, Kirsty J.; Nattkemper, Tim W.; Ruhl, Henry A.. 2016 Comparison of image annotation data generated by multiple investigators for benthic ecology. Marine Ecology Progress Series, 552. 61-70.10.3354/meps11775
Durden, Jennifer M.; Bett, Brian J.; Horton, Tammy; Serpell-Stevens, Amanda; Morris, Kirsty J.; Billett, David S.M.; Ruhl, Henry A.. 2016 Improving the estimation of deep-sea megabenthos biomass: dimension to wet weight conversions for abyssal invertebrates. Marine Ecology Progress Series, 552. 71-79.10.3354/meps11769
Dunlop, Katherine M.; van Oevelen, Dick; Ruhl, Henry A.; Huffard, Christine L.; Kuhnz, Linda A.; Smith, Kenneth L.. 2016 Carbon cycling in the deep eastern North Pacific benthic food web: Investigating the effect of organic carbon input. Limnology and Oceanography, 61 (6). 1956-1968.10.1002/lno.10345
Best, M.M.R.; Favali, P.; Beranzoli, L.; Blandin, J.; Cagatay, N.M.; Cannat, M.; Dañobeitia, J.J.; Delory, E.; de Miranda, J.M.A.; Del Rio Fernandez, J.; de Stigter, H.; Gillooly, M.; Grant, F.; Hall, P.O.J.; Hartman, S.; Hernandez-Brito, J.; Lanteri, N.; Mienert, J.; Oaie, G.; Piera, J.; Radulescu, V.; Rolin, J.-F.; Ruhl, H.A.; Waldmann, C.. 2016 The EMSO-ERIC Pan-European Consortium: data benefits and lessons learned as the legal entity forms. Marine Technology Society Journal, 50 (3). 8-15.
Cristini, Luisa; Lampitt, Richard S.; Cardin, Vanessa; Delory, Eric; Haugan, Peter; O'Neill, Nick; Petihakis, George; Ruhl, Henry A.. 2016 Cost and value of multidisciplinary fixed-point ocean observatories. Marine Policy, 71. 138-146.10.1016/j.marpol.2016.05.029
Lebrato, M.; Andersson, A.J.; Ries, J.B.; Aronson, R.B.; Lamare, L.B.; Koeve, W.; Oschlies, A.; Iglesias-Rodriguez, M.D.; Thatje, S.; Amsler, M.; Vos, S.; Jones, D.A.B.; Ruhl, H.A.; Gates, A.R.; McClintock, J.B.. 2016 Benthic marine calcifiers coexist with CaCO3-undersaturated seawater worldwide. Global Biogeochemical Cycles, 30 (7). 10.1002/2015GB005260
Morris, Kirsty J.; Bett, Brian J.; Durden, Jennifer M.; Benoist, Noelie M.A.; Huvenne, Veerle A.I.; Jones, Daniel O.B.; Robert, Katleen; Ichino, Matteo; Wolff, George A.; Ruhl, Henry A.. 2016 Landscape-scale spatial heterogeneity in phytodetrital cover and megafauna biomass in the abyss links to modest topographic variation. Scientific Reports, 6. 34080.10.1038/srep34080
Laguionie-Marchais, Claire; Paterson, Gordon L.J.; Bett, Brian J.; Smith, Kenneth L.; Ruhl, Henry A.. 2016 Inter-annual species-level variations in an abyssal polychaete assemblage (Sta. M, NE Pacific, 4000 m). Progress in Oceanography, 140. 43-53.10.1016/j.pocean.2015.10.006
Dunlop, Katherine M.; Kuhnz, Linda A.; Ruhl, Henry A.; Huffard, Christine L.; Caress, David W.; Henthorn, Richard G.; Hobson, Brett W.; McGill, Paul; Smith, Kenneth L.. 2015 An evaluation of deep-sea benthic megafauna length measurements obtained with laser and stereo camera methods. Deep Sea Research Part I: Oceanographic Research Papers, 96. 38-48.10.1016/j.dsr.2014.11.003
Ichino, Matteo C.; Clark, Malcolm R.; Drazen, Jeffery C.; Jamieson, Alan; Jones, Daniel O.B.; Martin, Adrian P.; Rowden, Ashley A.; Shank, Timothy M.; Yancey, Paul H.; Ruhl, Henry A.. 2015 The distribution of benthic biomass in hadal trenches: a modelling approach to investigate the effect of vertical and lateral organic matter transport to the seafloor. Deep Sea Research Part I: Oceanographic Research Papers, 100. 21-33.10.1016/j.dsr.2015.01.010
Laguionie-Marchais, C.; Kuhnz, L.A.; Huffard, C.L.; Ruhl, H.A.; Smith, K.L.. 2015 Spatial and temporal variation in sponge spicule patches at Station M, northeast Pacific. Marine Biology, 162 (3). 617-624.10.1007/s00227-014-2609-1
Main, C.E.; Ruhl, H.A.; Jones, D.O.B.; Yool, A.; Thornton, B.; Mayor, D.J.. 2015 Hydrocarbon contamination affects deep-sea benthic oxygen uptake and microbial community composition. Deep Sea Research Part I: Oceanographic Research Papers, 100. 79-87.10.1016/j.dsr.2014.12.008
Durden, Jennifer M.; Bett, Brian J.; Jones, Daniel O.B.; Huvenne, Veerle A.I.; Ruhl, Henry A.. 2015 Abyssal hills - hidden source of increased habitat heterogeneity, benthic megafaunal biomass and diversity in the deep sea. Progress in Oceanography, 137 (A). 209-218.10.1016/j.pocean.2015.06.006
Aguzzi, J.; Doya, C.; Tecchio, S.; De Leo, F.C.; Azzurro, E.; Costa, C.; Sbragaglia, V.; Del Río, J.; Navarro, J.; Ruhl, H.A.; Company, J.B.; Favali, P.; Purser, A.; Thomsen, L.; Catalán, I.A.. 2015 Coastal observatories for monitoring of fish behaviour and their responses to environmental changes. Reviews in Fish Biology and Fisheries, 25 (3). 463-483.10.1007/s11160-015-9387-9
Wynn, Russell B.; Huvenne, Veerle A.I.; Le Bas, Timothy P.; Murton, Bramley J.; Connelly, Douglas P.; Bett, Brian J.; Ruhl, Henry A.; Morris, Kirsty J.; Peakall, Jeffrey; Parsons, Daniel R.; Sumner, Esther J.; Darby, Stephen E.; Dorrell, Robert M.; Hunt, James E.. 2014 Autonomous Underwater Vehicles (AUVs): their past, present and future contributions to the advancement of marine geoscience. Marine Geology, 352. 451-468.10.1016/j.margeo.2014.03.012
Morris, Kirsty J.; Bett, Brian J.; Durden, Jennifer M.; Huvenne, Veerle A.I.; Milligan, Rosanna; Jones, Daniel O.B.; McPhail, Stephen; Robert, Katleen; Bailey, David M.; Ruhl, Henry A.. 2014 A new method for ecological surveying of the abyss using autonomous underwater vehicle photography. Limnology and Oceanography: Methods, 12 (11). 795-809.10.4319/lom.2014.12.795
Jones, Daniel O.B.; Yool, Andrew; Wei, Chih-Lin; Henson, Stephanie A.; Ruhl, Henry A.; Watson, Reg A.; Gehlen, Marion. 2014 Global reductions in seafloor biomass in response to climate change. Global Change Biology, 20 (6). 1861-1872.10.1111/gcb.12480
Best, M.; Favali, P.; Beranzoli, L.; Cannat, M.; Cagatay, N.; Dañobeitia, J.J.; Delory, E.; de Stigter, B.; Ferré, B.; Gillooly, M.; Grant, F.; Hall, P.O.J.; Lykousis, V.; Mienert, J.; de Miranda, J.M.A.; Oaie, G.; Radulescu, V.; Rolin, J.-F.; Ruhl, H.; Waldmann, C.. 2014 EMSO: A Distributed Infrastructure for Addressing Geohazards and Global Ocean Change. Oceanography, 27 (2). 167-169.10.5670/oceanog.2014.52
Kuhnz, Linda A.; Ruhl, Henry A.; Huffard, Christine L.; Smith, Kenneth L.. 2014 Rapid changes and long-term cycles in the benthic megafaunal community observed over 24years in the abyssal northeast Pacific. Progress in Oceanography, 124. 1-11.10.1016/j.pocean.2014.04.007
Ruhl, Henry A.; Bett, Brian J.; Hughes, Sarah J.M.; Alt, Claudia J.S.; Ross, Elizabeth J.; Lampitt, Richard S.; Pebody, Corinne A.; Smith, Kenneth L.; Billett, David S.M.. 2013 Links between deep-sea respiration and community dynamics. Ecology, 95 (6). 1651-1662.10.1890/13-0675.1
Laguionie-Marchais, C.; Billett, D.S.M.; Paterson, G.L.D.; Ruhl, H.A.; Soto, E.H.; Smith, K.L.; Thatje, S.. 2013 Inter-annual dynamics of abyssal polychaete communities in the North East Pacific and North East Atlantic—A family-level study. Deep Sea Research Part I: Oceanographic Research Papers, 75. 175-186.10.1016/j.dsr.2012.12.007
Wilson, S.E.; Ruhl, H.A.; Smith, K.L.. 2013 Zooplankton fecal pellet flux in the abyssal northeast Pacific: A 15 year time-series study. Limnology and Oceanography, 58 (3). 881-892.10.4319/lo.2013.58.3.0881
Mora, Camilo; Wei, Chih-Lin; Rollo, Audrey; Amaro, Teresa; Baco, Amy R.; Billett, David; Bopp, Laurent; Chen, Qi; Collier, Mark; Danovaro, Roberto; Gooday, Andrew J.; Grupe, Benjamin M.; Halloran, Paul R.; Ingels, Jeroen; Jones, Daniel O.B.; Levin, Lisa A.; Nakano, Hideyuki; Norling, Karl; Ramirez-Llodra, Eva; Rex, Michael; Ruhl, Henry A.; Smith, Craig R.; Sweetman, Andrew K.; Thurber, Andrew R.; Tjiputra, Jerry F.; Usseglio, Paolo; Watling, Les; Wu, Tongwen; Yasuhara, Moriaki. 2013 Biotic and Human Vulnerability to Projected Changes in Ocean Biogeochemistry over the 21st Century. PLoS Biology, 11 (10). e1001682.10.1371/journal.pbio.1001682
Jeffreys, Rachel M.; Burke, Ciara; Jamieson, Alan J.; Narayanaswamy, Bhavani E.; Ruhl, Henry A.; Smith, Kenneth L.; Witte, Ursula. 2013 Feeding Preferences of Abyssal Macrofauna Inferred from In Situ Pulse Chase Experiments. PLoS ONE, 8 (11). e80510.10.1371/journal.pone.0080510
Smith, K.L.; Ruhl, H.A.; Kahru, M.; Huffard, C.L.; Sherman, A.D.. 2013 Deep ocean communities impacted by changing climate over 24 y in the abyssal northeast Pacific Ocean. Proceedings of the National Academy of Sciences, 110 (49). 19838-19841.10.1073/pnas.1315447110
Vardaro, Michael F.; Bagley, Philip M.; Bailey, David M.; Bett, Brian J.; Jones, Daniel O.B.; Milligan, Rosanna J.; Priede, Imants G.; Risien, Craig M.; Rowe, Gilbert T.; Ruhl, Henry A.. 2013 A Southeast Atlantic deep-ocean observatory: first experiences and results. Limnology and Oceanography: Methods, 11. 304-315.10.4319/lom.2013.11.304
Hartman, S.E.; Lampitt, R.S.; Larkin, K.E.; Pagnani, M.; Campbell, J.; Gkritzalis, T.; Jiang, Z-P.; Pebody, C.A.; Ruhl, H.A.; Gooday, A.J.; Bett, B.J.; Billett, D.S.M.; Provost, P.; McLachlan, R.; Turton, J.D.; Lankester, S.. 2012 The Porcupine Abyssal Plain fixed-point sustained observatory (PAP-SO): variations and trends from the Northeast Atlantic fixed-point time-series. ICES Journal of Marine Science, 69 (5). 776-783.10.1093/icesjms/fss077
Drazen, Jeffrey C.; Bailey, David M.; Ruhl, Henry A.; Smith, Kenneth L.. 2012 The Role of Carrion Supply in the Abundance of Deep-Water Fish off California. PLoS ONE, 7 (11). e49332.10.1371/journal.pone.0049332
Kahn, Amanda S.; Ruhl, Henry A.; Smith, Kenneth L.. 2012 Temporal changes in deep-sea sponge populations are correlated to changes in surface climate and food supply. Deep Sea Research Part I Oceanographic Research Papers, 70. 36-41.10.1016/j.dsr.2012.08.001
Moeseneder, M.M.; Smith, K.L.; Ruhl, H.A.; Jones, D.O.B.; Witte, U.; Prosser, J.I.. 2012 Temporal and depth-related differences in prokaryotic communities in abyssal sediments associated with particulate organic carbon flux. Deep Sea Research Part I Oceanographic Research Papers, 70. 26-35.10.1016/j.dsr.2012.07.010
Vernet, Maria; Smith, Kenneth; Cefarelli, Adrián; Helly, John; Kaufmann, Ronald; Lin, Hai; Long, David; Murray, Alison; Robison, Bruce; Ruhl, Henry; Shaw, Timothy; Sherman, Alana; Sprintall, Janet. 2012 Islands of Ice: Influence of Free-Drifting Antarctic Icebergs on Pelagic Marine Ecosystems. Oceanography, 25 (3). 38-39.10.5670/oceanog.2012.72
Hughes, Sarah Jane Murty; Ruhl, Henry A.; Hawkins, Lawrence E.; Hauton, Chris; Boorman, Ben; Billett, David S.M.. 2011 Deep-sea echinoderm oxygen consumption rates and an interclass comparison of metabolic rates in Asteroidea, Crinoidea, Echinoidea, Holothuroidea and Ophiuroidea. The Journal of Experimental Biology, 214. 2512-2521.10.1242/jeb.055954
Ruhl, Henry A.; André, Michel; Beranzoli, Laura; Ça?atay, M. Namik; Colaço, Ana; Cannat, Mathilde; Dañobeitia, Juanjo J.; Favali, Paolo; Géli, Louis; Gillooly, Michael; Greinert, Jens; Hall, Per O.J.; Huber, Robert; Karstensen, Johannes; Lampitt, Richard S.; Larkin, Kate E.; Lykousis, Vasilios; Mienert, Jürgen; Miguel Miranda, J.; Person, Roland; Priede, Imants G.; Puillat, Ingrid; Thomsen, Laurenz; Waldmann, Christoph. 2011 Societal need for improved understanding of climate change, anthropogenic impacts, and geo-hazard warning drive development of ocean observatories in European Seas. Progress In Oceanography, 91 (1). 1-33.10.1016/j.pocean.2011.05.001
Ruhl, Henry A.; Ellena, Jacob A.; Wilson, Robert C.; Helly, John. 2011 Seabird aggregation around free-drifting icebergs in the northwest Weddell and Scotia Seas. Deep Sea Research Part II: Topical Studies in Oceanography, 58 (11-12). 1497-1504.10.1016/j.dsr2.2010.11.028
Ruhl, Henry A.; Priede, Imants G.. 2011 Open up monitoring of deep-sea drilling. Nature, 473 (7346). p154.10.1038/473154b
Ruhl, Henry A.; Rybicki, Nancy B.. 2010 Long-term reductions in anthropogenic nutrients link to improvements in Chesapeake Bay habitat. Proceedings of the National Academy of Sciences, 107 (38). 16566-16570.10.1073/pnas.1003590107
Smith, K.L.; Ruhl, H.A.; Bett, B.J.; Billett, D.S.M.; Lampitt, R.S.; Kaufmann, R.S.. 2009 Climate, carbon cycling and deep-ocean ecosystems. Proceedings of the National Academy of Sciences of the USA, 106. 19211-19218.10.1073/pnas.0908322106
Vardaro, Michael F.; Ruhl, Henry A.; Smith, Kenneth L.. 2009 Climate variation, carbon flux, and bioturbation in the abyssal North Pacific. Limnology and Oceanography, 54 (6). 2081-2088.
Holland, Nicholas D.; Jones, William J.; Ellena, Jacob; Ruhl, Henry A.; Smith, Kenneth L.. 2009 A new deep-sea species of epibenthic acorn worm (Hemichordata, Enteropneusta). Zoosystema, 321 (2). 333-346.
Ruhl, Henry A.; Ellena, Jacob A.; Smith, Kenneth L.. 2008 Connections between climate, food limitation, and carbon cycling in abyssal sediment communities. Proceedings of the National Academy of Sciences of the USA, 105 (44). 17006-17011.10.1073/pnas.0803898105
Smith, K.L.; Ruhl, H.A.; Kaufmann, R.S.; Kahru, M.. 2008 Tracing abyssal food supply back to upper-ocean processes over a 17-year time series in the northeast Pacific. Limnology and Oceanography, 53 (6). 2655-2667.
Booth, J. Ashley T.; Ruhl, Henry A.; Lovell, Lawrence L.; Bailey, David M.; Smith, Kenneth L.. 2008 Size–frequency dynamics of NE Pacific abyssal ophiuroids (Echinodermata: Ophiuroidea). Marine Biology, 154 (6). 933-941.10.1007/s00227-008-0982-3
Smith, K.L.; Robison, B.H.; Helly, J.J.; Kaufmann, R.S.; Ruhl, H.A.; Shaw, T.; Twining, B.S.; Vernet, M.. 2007 Free-drifting icebergs: hotspots of chemical and biological enrichment in the Weddell Sea. Science, 317 (5837). 478-482.10.1126/science.1142834
Smith, K.L.; Baldwin, R.J.; Ruhl, H.A.; Kahru, M.; Mitchell, B.G.; Kaufmann, R.S.. 2006 Climate effect on food supply to depths greater than 4,000 meters in the northeast Pacific. Limnology and Oceanography, 51 (1). 166-176.
Bailey, D.M.; Ruhl, H.A.; Smith, K.L.. 2006 Long-term changes in benthopelagic fish abundance in the abyssal N.E. Pacific Ocean. Ecology, 87 (3). 549-555.10.1890/04-1832
Smith, K.L.; Holland, Nicholas D.; Ruhl, Henry A.. 2005 Enteropneust production of spiral fecal trails on the deep-sea floor observed with time-lapse photography. Deep-Sea Research I, 52 (7). 1228-1240.10.1016/j.dsr.2005.02.004
Ruhl, Henry A.; Smith, Kenneth L.. 2004 Shifts in deep-sea community structure linked to climate and food supply. Science, 305 (5683). 513-515.10.1126/science.1099759
Ruhl, Henry A.; Hastings, Philip A.; Zarubick, Lisa A.; Jensen, Rachelle M.; Zdzitowiecki, Krzysztof. 2003 Fish populations of Port Foster, Deception Island, Antarctica and vicinity. Deep-Sea Research II, 50 (10-11). 1843-1858.10.1016/S0967-0645(03)00094-8
Kendall, Katrina A.; Ruhl, Henry A.; Wilson, Robert C.. 2003 Distribution and abundance of marine bird and pinniped populations within Port Foster, Deception Island, Antarctica. Deep-Sea Research II, 50 (10-11). 1873-1888.10.1016/S0967-0645(03)00096-1
Rybicki, Nancy B.; McFarland, Dwilette G.; Ruhl, Henry A.; Reel, Justin T.; Barko, John W.. 2001 Investigations of the availability and survival of submersed aquatic vegetation propagules in the tidal Potomac River. Estuaries, 24 (3). 407-424.10.2307/1353242

Book section

Matabos, M.; Best, M.; Blandin, J.; Hoeberechts, M.; Juniper, S.K.; Pirenne, B.; Robert, K.; Ruhl, H.; Sarrazin, J.; Vardaro, M.. 2016 Seafloor observatories. In: Clark, M.; Consalvey, M.; Rowden, A.A., (eds.)Biological Sampling in the Deep-Sea. Chichester, John Wiley & Sons, 306-337, 472pp.
Durden, Jennifer M.; Schoening, Timm; Althaus, Franziska; Friedman, Ariell; Garcia, Rafael; Glover, Adrian G.; Greinert, Jens; Jacobsen Stout, Nancy; Jones, Daniel O.B.; Jordt, Anne; Kaeli, Jeffrey W.; Köser, Kevin; Kuhnz, Linda A.; Lindsay, Dhugal; Morris, Kirsty J.; Nattkemper, Tim W.; Osterloff, Jonas; Ruhl, Henry A.; Singh, Hanumant; Tran, Maggie; Bett, Brian J.. 2016 Perspectives in visual imaging for marine biology and ecology: from acquisition to understanding. In: Hughes, R.N.; Hughes, D.J.; Smith, I.P.; Dale, A.C., (eds.)Oceanography and Marine Biology: An Annual Review, Vol. 54. Boca Raton, FL, CRC Press, 1-72, 470pp.
Bagley, P.M.; Smith, K.L.; Bett, B.J.; Priede, I.G.; Rowe, G.; Clarke, J.; Walls, A.; Ruhl, H.A.; Bailey, D.M.. 2011 The DELOS project, long-term observatories in an oil field environment in the Tropical Atlantic Ocean off Angola, West Africa. In: Favali, P.; de Santis, A.; Beranzoli, L., (eds.)Seafloor observatories: A new vision of the earth from the abyss. Berlin, Springer-Praxis, Chap.11, 500pp. (In Press)
Person, R.; Rolin, J.F.; Andre, M.; Auffret, Y.; Blandin, J.; Beranzoli, L.; Cagatay, N.; Cannat, M.; Colaco, A.; Danobeitia, J.J.; Delory, E.; Drogou, J.F.; Fahrbach, E.; Favali, P.; Gifci, G.; Geli, L.; Gillooly, M.; Grant, F.; Henry, P.; Hall, P.; Huber, R.; Karstensen, J.; Klages, M.; Lampitt, R.; Lykousis, A.; Mienert, J.; Miranda, J.M.; Priede, I.E.; Ruhl, H.; Thomsen, L.; Tselepides, A.; Waldman, C.; Wenzhoefer, F.; Wright, I.C.. 2011 ESONET NoE: An European initiative to develop and integrate the scientific European community using deep-sea observatories. In: Favali, P.; de Santis, A.; Beranzoli, L., (eds.)Seafloor observatories: A new vision of the earth from the abyss. Berlin, Springer-Praxis, Chap.14, 500pp. (In Press)
Larkin, K.E.; Ruhl, H.A.; Bagley, P.; Benn, A.; Bett, B.J.; Billett, D.S.M.; Boetius, A.; Chevaldonné, P.; Colaço, A.; Copley, J.; Danovaro, R.; Escobar-Briones, E.; Glover, A.; Gooday, A.J.; Hughes, J.A.; Kalogeropoulou, V.; Kitazato, H.; Kelly-Gerreyn, B.A.; Klages, M.; Lampadariou, N.; Lejeusne, C.; Perez, T.; Priede, I.G.; Rogers, A.; Sarradin, P.M.; Sarrazin, J.; Soltwedel, T.; Soto, E.H.; Thatje, S.; Tselepides, A.; van den Hove, S.; Tyler, P.A.; Vanreusel, A.; Wenzhöfer, F.. 2010 Benthic biology time-series in the deep sea: indicators of change. In: Hall, J.; Harrison, D.E.; Stammer, D., (eds.)Proceedings of OceanObs’09: Sustained Ocean Observations and Information for Society, Vol. 2. Noordwijk, The Netherlands, European Space Agency, 599-615. (ESA Special Publication, WPP-306).
Glover, A.G.; Gooday, A.J.; Bailey, D.M.; Billett, D.S.M.; Chevaldonne, P.; Colaco, A.; Copley, J.; Cuvelier, D.; Desbruyeres, D.; Kalogeropoulou, V.; Klages, M.; Lampadariou, N.; Lejeunsne, C.; Mestre, N.C.; Paterson, G.L.J.; Perez, T.; Ruhl, Henry A.; Sarrazin, J.; Soltwedel, T.; Soto, E.H.; Thatje, S.; Tselepides, A.; Van Gaever, S.; Vanreusel, A.. 2010 Temporal change in deep-sea benthic ecosystems: a review of the evidence from recent time-series studies. In: Lesser, M., (ed.)Advances in Marine Biology, Vol. 58. London, UK, Academic Press, 1-95, 352pp.


Ruhl, H.A.; et al, .. 2016 RRS Discovery Cruise DY034, 06 Aug - 01 Sep 2015, Southampton to Southampton. Shelf sea biogeochemistry. Southampton, National Oceanography Centre, 121pp. (National Oceanography Centre Cruise Report 39)
Ruhl, H.A.; et al, .. 2015 RRS Discovery Cruise DY008, 18 Mar - 13 Apr 2014, Southampton to Southampton. Shelf sea biogeochemistry. Southampton, National Oceanography Centre, 133pp. (National Oceanography Centre Cruise Report 33)
Ruhl, H.A.; et al, .. 2012 RRS James Cook Cruise 62, 24 Jul-29 Aug 2011. Porcupine Abyssal Plain – sustained observatory research. Southampton, UK, National Oceanography Centre Southampton, 119pp. (National Oceanography Centre Cruise Report 12)

Conference item

Morris, Kirsty; Jones, Daniel; Bett, Brian; Billett, David; Huvenne, Veerle; McPhail, Stephen; Ruhl, Henry. 2013 Autonomous Ecological Surveying of the Abyss (AESA):Towards a landscape scale perspective in deep-sea ecology. [Poster] In: UK-IMON International Workshop on New Monitoring Technologies:A Workshop to Prioritise Future Investment, Southampton, 10-12 Sep 2013. Southampton, National Oceanography Centre. (Unpublished)